Martin - these transistors were part of a long series of transistors that were all very similar, mainly just with different cases and pinouts. Elektor magazine used to have a page in every issue called TUPTUNDUGDUS - which stood forTransistor, Universal, NPNTransistor, Universal, PNP
Diode, Universal, Germanium
Diode, Universal, Silicon
They used to specify the above as components in many projects, which made life very simple if you had a reasonable junk box! In the page they gave lists of parts which met the specifications for each component. They don't do this any more (not a lot of projects use transistors now

but it's still just as valid. I was fortunate enough to find a copy of their page on the net here. I'd think that for most purposes you'd be able to use this as a substitution list for this and other projects. Many people have linked to it - just put TUPTUNDUGDUS into google and you'll find one. I keep a copy ready to hand for just this purpose. Bear in mind that cases and pinouts are different, so you may have to work out your lead orientation if you use a different transistor, but they'll almost always work. There is a list of pinouts down the right hand side, along with remarks in which your transistors are noted as having an Ic max of 200mA, which is greater than their minimum spec for TUNs and TUPs. If this is likely to be an issue in your circuit then select parts with similar limits. But for Audio applications that's not likely.In rare (and I would say very rare) cases, a circuit may depend on a part having a certain gain or power or current or frequency response, but for Audio circuits this would not usually be a problem. And lines like the BC547-8-9 and BC557-8-9 often came with an A,B or C suffix to indicate progressively higher gain types - so use those if you know a high gain transistor is needed.
Hope this helps // David